True love can last a lifetime - and that is evident in both humans and animals.

Earlier this week, the Los Angeles Zoo announced that animal care staff had made the difficult decision to put two 21-year-old lions to sleep at the same time, after six years of being lovingly inseparable from each other.

Writing on the official LA Zoo and Botanical Gardens Facebook page, a spokesperson provided the following statement: "It is with a heavy heart that we announce the loss of our African lion pair, Hubert and Kalisa.

"Animal care and health staff made the difficult decision to humanely euthanize the 21-year-old lions today due to their declining health and age-related illnesses that had diminished their quality of life."

CEO & Zoo Director Denise Verret said of the lions' passing: "Hubert and Kalisa are an iconic part of the LA Zoo experience, and our staff and guests have been touched by their loyal companionship."

"These affectionate companions came to the LA Zoo six years ago, and they quickly charmed themselves into our hearts as we observed their magnificent beauty and unique bond. It was often said, you don’t see Kalisa without Hubert being close by.

"So, while it is truly heart-wrenching that we had to say goodbye to this iconic pair, we can take comfort in knowing they left together. These lions will remain a positive part of our history, and they will be greatly missed."

Verret added: "I have to commend our animal care and veterinary staff for the great care they’ve given this pair, a couple who lived longer than most lions do in human care and the wild."

The Facebook statement continued: "With an average life expectancy of mid-teens and about 17 years in Zoos, Hubert and Kalisa were considered elderly when they arrived at the LA Zoo in 2014 from the Woodland Park Zoo, where they had first bonded.

"They quickly became favorites among L.A. Zoo guests and staff and were known for their frequent cuddles and nuzzles.

"Please join us in honoring Hubert and Kalisa and their legacy by sharing some of your favorite memories of them here or by tagging LA Zoo."

LA Zoo spokesperson Beth Schaefer told the Los Angeles Times:

"These lions were charismatic both together as partners and separately, but they were hardly ever apart from one another.

"Their undivided attention was always on the other as they rested together, cuddled and nuzzled often."

Our hearts go out to all the keepers and visitors of the LA Zoo.

True love can last a lifetime - and that is evident in both humans and animals.

Earlier this week, the Los Angeles Zoo announced that animal care staff had made the difficult decision to put two 21-year-old lions to sleep at the same time, after six years of being lovingly inseparable from each other.

Writing on the official LA Zoo and Botanical Gardens Facebook page, a spokesperson provided the following statement: "It is with a heavy heart that we announce the loss of our African lion pair, Hubert and Kalisa.

"Animal care and health staff made the difficult decision to humanely euthanize the 21-year-old lions today due to their declining health and age-related illnesses that had diminished their quality of life."

CEO & Zoo Director Denise Verret said of the lions' passing: "Hubert and Kalisa are an iconic part of the LA Zoo experience, and our staff and guests have been touched by their loyal companionship."

"These affectionate companions came to the LA Zoo six years ago, and they quickly charmed themselves into our hearts as we observed their magnificent beauty and unique bond. It was often said, you don’t see Kalisa without Hubert being close by.

"So, while it is truly heart-wrenching that we had to say goodbye to this iconic pair, we can take comfort in knowing they left together. These lions will remain a positive part of our history, and they will be greatly missed."

Verret added: "I have to commend our animal care and veterinary staff for the great care they’ve given this pair, a couple who lived longer than most lions do in human care and the wild."

The Facebook statement continued: "With an average life expectancy of mid-teens and about 17 years in Zoos, Hubert and Kalisa were considered elderly when they arrived at the LA Zoo in 2014 from the Woodland Park Zoo, where they had first bonded.

"They quickly became favorites among L.A. Zoo guests and staff and were known for their frequent cuddles and nuzzles.

"Please join us in honoring Hubert and Kalisa and their legacy by sharing some of your favorite memories of them here or by tagging LA Zoo."

LA Zoo spokesperson Beth Schaefer told the Los Angeles Times:

"These lions were charismatic both together as partners and separately, but they were hardly ever apart from one another.

"Their undivided attention was always on the other as they rested together, cuddled and nuzzled often."

Our hearts go out to all the keepers and visitors of the LA Zoo.