The marine animal community and its legion of supporters are in mourning as Lolita, an orca who had spent over half a century in captivity at the Miami Seaquarium, died at the estimated age of 57.
She passed away from suspected kidney disease, becoming a symbol of the complex ethical debates surrounding marine life in captivity, per Sky News.
Also known by the names Tokitae or Toki, she held the distinction of being the world's second-oldest known killer whale. Her death last Friday struck an emotional chord as it happened just as preparations were underway to return her to the ocean.
The Miami Seaquarium revealed in a statement that Lolita had "started exhibiting serious signs of discomfort" recently. Despite being "treated immediately and aggressively," the iconic whale died from what is believed to be "a renal condition."

The staff lauded her as "an inspiration" and a "beautiful spirit," giving a nod to the Lummi Nation, a Native American tribe based in Washington state, who considered her as part of their extended family.
For decades, Lolita had been a star attraction, performing tricks for Miami's crowds after being captured in 1970 at the tender age of four. In a poignant contrast, the orca believed to be her mother, Ocean Sun, swims free in the waters between Washington state and Canada and is currently in her 90s.
Lolita's life had been the focus of relentless campaigns by animal rights activists, pressing for her release from her 80ft by 35ft tank. She officially retired from public performances last spring, fulfilling a condition for the Seaquarium's new exhibitor's license from the US Department of Agriculture.
Earlier this year, a ray of hope emerged for Lolita. The park's new owner, The Dolphin Company, and the non-profit Friends of Toki disclosed plans, backed by Indianapolis Colts owner Jim Irsay, to move her to a more natural sea pen in the Pacific Northwest.
Responding to the tragic news, Irsay said, "I am heartbroken that Toki has left us. Her story captured my heart, just as it did millions of others." Eduardo Albor, President of The Dolphin Company, expressed his devastation on social media, lauding the efforts of care team led by Mike Partica and Dr. Reiderson as "true heroes."
Tony Hillaire, chairman of the Lummi Nation, summed up the collective grief: "The Lummi Nation is saddened by the news that our beloved Orca relative has passed away at the estimated age of 57 years old. Our hearts are with all those impacted by this news; our hearts are with her family."
While new upgrades had been implemented in recent months to improve her living conditions, Lolita's much-anticipated freedom still faced bureaucratic obstacles. Federal and state regulators would have needed to green-light plans for her relocation, a process that could have taken years.
As the world reflects on Lolita's life and what could have been, her legacy will undoubtedly fuel ongoing debates about marine captivity and conservation. The orca who captured the hearts of millions will long be remembered, even as questions about ethical treatment of marine life continue to make waves.